University of Toledo running backs coach Louis Ayeni is leaving to take the same job at Iowa State.
Ayeni, who held the title as associate head coach, produced a 1000-yard back in all four of his seasons with the Rockets.
Iowa State fired running backs coach Kenith Pope last month.
Toledo, after experiencing zero staff turnover last offseason, has lost three assistants this year. Defensive coordinator Tom Matukewicz left to be head coach at Southeast Missouri State and took with him safeties coach Bryce Saia.